Whether to enable special handling of the IPv4 default route.
If enabled, the IPv4 default route from wireguard.peer-routes will be placed to a dedicated routing-table and two policy routing rules will be added. The fwmark number is also used as routing-table for the default-route, and if fwmark is zero, an unused fwmark/table is chosen automatically. This corresponds to what wg-quick does with Table=auto and what WireGuard calls "Improved Rule-based Routing".
Note that for this automatism to work, you usually don't want to set ipv4.gateway, because that will result in a conflicting default route.
Leaving this at the default will enable this option automatically if ipv4.never-default is not set and there are any peers that use a default-route as allowed-ips. Since this automatism only makes sense if you also have a peer with an /0 allowed-ips, it is usually not necessary to enable this explicitly. However, you can disable it if you want to configure your own routing and rules.